From A. R. Wallace   21 July 1875

The Dell, Grays, Essex

July 21st. 1875

Dear Darwin

Many thanks for your kindness in sending me a copy of your new book.1 Being very busy I have only had time to dip into it yet. The account of Utricularia is most marvellous & quite new to me. I am rather surprised that you do not make any remarks on the origin of these extraordinary contrivances for capturing insects.2 Did you think they were too obvious? I dare say there is no difficulty, but I feel sure they will be seized on as inexplicable by Nat. Select. & your silence on the point will be held to show that you consider them so! The contrivance in Utricularia & Dionæa, & in fact in Drosera too, seems fully as great & complex as in Orchids, but there is not the same motive force.3 Fertilization & cross fertilization are important ends enough to lead to any modification,—but can we suppose mere nourishment to be so important, seeing that it is so easily & almost universally obtained by extension of roots & leaves. Here are plants which lose their roots & leaves to acquire the same results by infinitely complex modes! What a wonderfull & long continued series of variations must have led up to the perfect “trap” in Utricularia, while at any stage of the process the same end might have been gained by a little more development of roots & leaves, as in 9999 plants out of 10,000!

Is this an imaginary difficulty or do you mean to deal with it in future editions of the “Origin”?4

Believe me | Yours very faithfully| Alfred R. Wallace

Wallace’s name is on the presentation list for Insectivorous plants (Appendix IV).
In Insectivorous plants, pp. 404–13, CD described the function of the small bladders of Utricularia (bladderwort) in capturing and absorbing insects.
Dionaea (Venus fly trap) and Drosera (sundew) were the principle genera discussed in Insectivorous plants. Wallace alludes to Orchids.
Apart from a reprint issued in 1876 with minor corrections, there were no further revisions of Origin published in CD’s lifetime (see Freeman 1977, pp. 80–1).

Please cite as “DCP-LETT-10085,” in Ɛpsilon: The Charles Darwin Collection accessed on 5 June 2025, https://epsilon.ac.uk/view/dcp-data/letters/DCP-LETT-10085