From H. C. Watson   10 March 1857

Thames Ditton

March 10th/57

My dear Sir

I am not quite sure of understanding your question about “variable genera”.1 To explain my uncertainty, I will endeavour to define or state the differences for choice.

1. Genera, of which the species are close, & difficult to distinguish by reason of their similarity;—but the species themselves not remarkably variable. Ex: Carex & Ranunculus (excluding Batrachium)

2. Genera, of which the quasi species are so close that it becomes highly difficult to say whether the genus is composed of a comparatively few extremely variable species, or of many very close species. Ex: Rubus & Hieracium.

3. Genera, the species of which are themselves so variable, & approximating, that it becomes difficult to say where one species ends & the next begins. Ex: Viola & Saxifraga, at least in certain sections or subgenera—

It seems to me that Dr. A. Gray may have inclined to the first, while you perhaps yes certainly intend 2 or 3. I will copy the three categories, that you maybe

CD annotations

crossed pencil
Top of first page: ‘Please return to me’ink; ‘1)’ink 2
CD had asked Watson to comment on the list of ‘protean’ genera (in which the species present a great amount of variation) included in the letter from Asa Gray, 16 February 1857. CD discussed protean genera in Origin, p. 46.
The annotations relate to a later occasion when CD forwarded Watson’s letter to Asa Gray. See letter to Asa Gray, [after 15 March 1857].

Please cite as “DCP-LETT-2063,” in Ɛpsilon: The Charles Darwin Collection accessed on 5 June 2025, https://epsilon.ac.uk/view/dcp-data/letters/DCP-LETT-2063