To G. H. K. Thwaites   29 December [1862]

Down. | Bromley. | Kent. S.E.

Dec. 29th

My dear Mr Thwaites

Can you give me any cases of what some gardeners call “Sports”, but what I shall call “bud-variations”, ie when a leaf-bud assumes suddenly a new character—like moss-rose on Provence &c.— I shd be very grateful for any authentic cases.—1 Why I more particularly ask you, is that Sir R. Schomburghk says that at St Domingo, introduced flowers from the warmer temperate regions were there particularly apt to sport in this manner.2

I wrote some time ago to you about Cinchona being dimorphic like Primula;3 I have now better reason to believe that this is the case, & I I have proof that some of these dimorphic plants are absolutely sterile with their own-form pollen.—4

You mentioned two Ceylon genera (I have your note but names forgotten; one was allied to Menyanthes)5   Would you have kindness to examine them & see that both forms produce pollen & if you will, compare size of pollen. Or if you prefer could you send me the two forms of these genera in a letter, well dried, but not much pressed & named & then I could compare by soaking the pollen. The flowers shd not be old.6

I believe that you will forgive me for troubling you, & I remain | Yours very sincerely | Ch. Darwin

I am hard at work at a book on “Variation under Domestication”.7

According to his ‘Journal’ (Correspondence vol. 10, Appendix II), CD began working on chapter 11 of Variation, ‘On bud-variation, and on certain anomalous modes of reproduction and variation’, on 21 December 1862.
Schomburgk 1858.
See letter to Asa Gray, 14 July [1862] and nn. 11 and 12.
CD had begun work on Variation, the first part of a projected three-part work on natural selection, in January 1860 (Correspondence vol. 8); it was not published until 1868.

Manuscript Alterations and Comments

3.4 of these genera] interl
3.5 much] interl

Please cite as “DCP-LETT-3880,” in Ɛpsilon: The Charles Darwin Collection accessed on 5 June 2025, https://epsilon.ac.uk/view/dcp-data/letters/DCP-LETT-3880