From H. W. Bates   20 April 1868

Royal Geographical Society | 15, Whitehall Place, S.W.

April 20 1868

My dear Mr Darwin

I have found amongst my old papers a rude drawing of the conspicuous caterpillar which you intend to mention so enclose it.1 I believe it is natural size; the black is not deep enough, otherwise I think the colour is accurate

The species of Æneas group of Papilio in which the sexes show no difference in colours or pattern are2

P. Triopas } males simply have deeper & richer ground colour than females
P. Chabrias
P. Montezuma
P. Panthonus
P. Ascanius — I see no difference in tint of ground colour.

P. Ascanius in both sexes is handsome; I think more conspicuous & gay in colouring than any of the other four species here mentioned. The whole Æneas group is remarkable for its rich, strongly-constrated3 & conspicuous colouring; but the rule is—less richness of tint in ♀ than in ♂, although the ♀ has often more variety of colour. It is very clear there is no adaptation of female colouring to disguise in this group.

The females of Argynnis Diana & A. Sagana are of olive-brown or blackish colours relieved by lighter marks.4 Regarded absolutely one cannot consider them less beautiful than their males; but there is no doubt, here, adaptation for purposes of disguise has taken place; Entomologists of the countries where they are found alone could say what they imitate; I have no idea myself; but A. Sagana ♀ is coloured much like many common butterflies of Adolias group found in some countries.5

Yours sincerely | H W Bates

A capital suggestion that of H. Doubleday!6

CD annotations

1.1 I have … accurate 1.3] crossed pencil
3.1 The females … countries. 3.6] crossed pencil
The enclosure has not been found, but in Descent 1: 416, CD gave Bates’s description of the caterpillar, ‘the most conspicuous … which he ever beheld’.
For more on the ‘Æneas group’, see the letter from H. W. Bates, 12 March 1868 and nn. 5 and 10. All the species mentioned in the list now belong to the genus Parides.
Bates evidently intended ‘strongly-contrasted’.
Argynnis diana is a synonym of Speyeria diana, the Diana fritillary; both it and Argynnis sagana are highly sexually dimorphic.
The genus Adolias is now Euthalia (barons).

Please cite as “DCP-LETT-6132,” in Ɛpsilon: The Charles Darwin Collection accessed on 5 June 2025, https://epsilon.ac.uk/view/dcp-data/letters/DCP-LETT-6132