To J. D. Hooker   10 November [1861]

Down Bromley Kent

Nov. 10th

My dear Hooker

For the love of God help me. I believe all my work (about a fortnight) is useless. Look at this accursed diagram of the Butterfly orchis, which I examined after writing to you yesterday when I thought all my work done.1 Some of The ducts of upper sepal & upper petal run to the wrong bundles or column.— I have seen no such case. This case apparently shows that not the least reliance can be placed on course of ducts. (I am sure of my facts).

There is great adhesion & extreme displacement of parts where the organs spring from top of ovarium. Asa Gray says ducts are very early developed, & it seems to me wonderful that they should pursue this course.— It may now be said that the lateral ducts in the Labellum running into the antero-lateral ovarian bundle is no argument that the Labellum consists of 3 organs blended together.—2

In desperation (& from the curious way the base of upper petals are soldered at basal edges) I fancied the real form of upper Sepal, upper petal, & lower sepal might be as represeted by red lines;3 & that there had been an incredible amount of splitting of sepals & petals & subsequent fusion.— This seems a monstrous notion; but I have just looked at Bauer’s drawing of allied Bonatea, & there is a degree of lobing of petals & sepals, which would account for anything.4 Now could you spare me a dry flower out of your Herbarium of Bonatea speciosa, that I might soak & look for ducts; if I cannot explain the case of Habenaria, all my work is smashed. I was a fool ever to touch orchids.—

Your affect. | C. Darwin

[Enclosure]

diagram
See the enclosure.
CD was attempting to trace the course of the spiral vessels in orchid flowers, from the ovarium to the various parts, in an attempt to establish the homologies of orchids (see letters to J. D. Hooker, 4 October [1861], 13 October [1861], 27 October [1861], and the first letter dated 1 November [1861]). In the account he gave of orchid homologies in Orchids, CD discussed the anomaly presented by Habenaria and the allied genus Bonatea, stating that it called into question his view that the labellum was always ‘an organ compounded of one petal and two petaloid stamens’ (Orchids, p. 303).
CD drew the outermost lines, representing the ‘real form’ of the sepal and petal parts, in red crayon. The diagram has been reduced to 60% of its original size.
Bauer 1830–8.

Manuscript Alterations and Comments

1.3 Some of] interl
1.4 bundles or] interl
1.6 (I am sure of my facts)] square brackets in MS
3.3 & that] after del ‘but’; ‘&’ over illeg
3.7 out of your Herbarium] interl
Diagram
(Dotted lines ducts)] square brackets in MS

Please cite as “DCP-LETT-3315,” in Ɛpsilon: The Charles Darwin Collection accessed on 5 June 2025, https://epsilon.ac.uk/view/dcp-data/letters/DCP-LETT-3315